Level 1
Frost Spines (Quest): Attack enemy heroes while affected by Tidal Burst 30 times.
Reward: Ranged attacks now slow enemies by 20% and extend Tidal Burst's duration by 1 second.
Harden (Trait): Switching to Myrmidon Stance grants 2 charges of block that reduce physical damage by 50%.
Swipe Master (Quest) Hit enemy heroes with Horseshoe Swipe or Double Swipe 50 times.
Reward: Increases Horseshoe Swipe range and damage by 20% and Double Swipe now causes Naj'entus to lunge at his target.
Level 4
Resilience (W): Increases tidal shield duration by 0.5 seconds each time Naj'entus is stunned, rooted or silenced.
Vicious Spine (W): Increases needle spine's splash damage by 30% and increases the area of effect by 20%.
Acid Membrane (E): The acid mortar's projectile bounces once if it does not hit an enemy.
Coagulating Slime (E): Vile slime also reduces attack speed by 25%.
Level 7
Shell Smash (Trait): Switching to Impaler Stance increases attack speed by 30% for 4 seconds.
Ram (Q): Horsheshoe swipe now knocks back the first enemy hero it hits and stuns them for 1 second.
Bubble Burst (W): 50% of the damaged absorbed by tidal shield is dealt to nearby enemies when the shield expires.
Level 10 (See Heroic Abilities)
Level 13
Double Back (Q): horseshoe swipe can be reactivated within 1.5 seconds to cause Naj'entus to jump backwards.
Ink Blast (Q): Tidal burst now blinds the target for 2 seconds.
Sludge Bomb (E): Vile slime now affects enemies in an area but cooldown is increased by 3 seconds.
Level 16
Water Spout (Q): Tidal Burst deals 75% increased damage at full health. This damage decreases down to zero at 75% health.
Iron Scale (W): Gain 30 armor for 3 seconds after tidal shield expires.
Impaling Spine (W): Needle spine roots the first target it hits for 2 seconds.
Acid Barrage (E): Acid mortar's mana cost and cooldown is refunded if it does not hit any enemies.
Level 20
Bed of Spines (See Heroic abilities)
Brine Tide (See Heroic abilities)
Leech Spine (Trait): Ranged attacks gain 30% life steal, Melee attacks gain 15% life steal.
Lacerating Shell (Trait): Double swipe now deals 9% of an enemy's health in damage. Every 3rd ranged attack now deals 50% increased damage.
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